Click on 2 adjacent quarks to try swapping their positions.
The swap will be successful if it results in a match of 3 or more quarks vertically or horizontally.
A failed swap has no consequences.
Matched quarks will disappear from the board. (If you've played Bejewelled, this should sound pretty familiar so far.)
The arrow direction on matched quarks indicates the direction surrounding quarks will fall to fill the space left behind.
Quarks only match if they have the same arrow direction and ring color.
The game ends after 5 successful swaps, at which point it will say you've won. Don't get too excited.
There are still a few bugs and unimplemented mechanics that might break the game sometimes, but it works ~90% of the time.